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Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Aves
Order: Passeriformes
Superfamily: Bombycilloidea
Family: Hylocitreidae
Fjeldsa, Ericson, Johannson, & Zuccon 2015
Genus: Hylocitrea
Mathews, 1925
H. bonensis
Binomial name
Hylocitrea bonensis
(Meyer & Wiglesworth, 1894)

The hylocitrea (Hylocitrea bonensis), also known as the yellow-flanked whistler or olive-flanked whistler, is a species of bird that is endemic to montane forests on the Indonesian island of Sulawesi.[3] It is monotypic within the genus Hylocitrea, and has traditionally been considered a member of the family Pachycephalidae, but recent genetic evidence suggests it should be placed in a monotypic subfamily of the family Bombycillidae,[4] or even its own family, Hylocitreidae.[1] A 2019 study found it to be a sister group to a clade containing the hypocolius (Hypocoliidae) and the extinct Hawaiian honeyeaters (Mohoidae), with the clade containing all three being a sister group to the silky-flycatchers (Ptiliogonatidae). The divergences forming these families occurred in the early Miocene, about 20-23 million years ago.[5]


  1. ^ a b Gill, F; D Donsker, eds. (2011). "Waxwings to Swallows". IOC World Bird Names (version 2.9). Archived from the original on July 24, 2011. Retrieved 17 August 2011.
  2. ^ BirdLife International (2016). "Hylocitrea bonensis". IUCN Red List of Threatened Species. 2016: e.T103867190A94017773. doi:10.2305/IUCN.UK.2016-3.RLTS.T103867190A94017773.en. Retrieved 16 November 2021.
  3. ^ Boles, W. E. (2007). Yellow-flanked Whistler (Hylocitrea bonensis). pp. 411 in: Del Hoyo, J.; Elliot, A., & Christie D. eds. (2007). Handbook of the Birds of the World. Vol. 12. Picathartes to Tits and Chickadees. Lynx Edicions, Barcelona. ISBN 978-84-96553-42-2
  4. ^ Spellman, G.A.; Cibois, A; Moyle, RG; Winker, K; Keith Barker, F; et al. (2008). "Clarifying the systematics of an enigmatic avian lineage: What is a bombycillid?". Molecular Phylogenetics and Evolution. 49 (3): 691–1044. doi:10.1016/j.ympev.2008.09.006. PMC 2627281. PMID 18824237.
  5. ^ Lerner, Heather R.L.; Meyer, Matthias; James, Helen F.; Hofreiter, Michael; Fleischer, Robert C. (2011-11-08). "Multilocus Resolution of Phylogeny and Timescale in the Extant Adaptive Radiation of Hawaiian Honeycreepers". Current Biology. 21 (21): 1838–1844. doi:10.1016/j.cub.2011.09.039. ISSN 0960-9822. PMID 22018543.
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