Symmetrodonta Temporal range: Early Cretaceous–Campanian PreꞒ Ꞓ O S D C P T J K Pg N Possible Paleocene ghost lineage
Fossil specimen of Zhangheotherium quinquecuspidens
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Clade:	Trechnotheria
Order:	†Symmetrodonta
Subgroups
†?Chronoperates †Spalacotheriidae †Zhangheotheriidae

Symmetrodonta is a group of Mesozoic mammals and mammal-like synapsids characterized by the triangular aspect of the molars when viewed from above, and the absence of a well-developed talonid. The traditional group of 'symmetrodonts' ranges in age from the latest Triassic to the Late Cretaceous, but most research in the last 20-30 years has concluded that they are not a true taxonomic group, but include several unrelated branches of the mammal tree. Despite this, the name is still used informally by some researchers for convenience, usually restricted to the spalacotheriids and zhangheotheriids.^[1][2]

There are some symmetrodonts with acutely-triangulated molar cusps (“acute-angled symmetrodonts”) that seem to form a true monophyletic group, and lasted from the Early Cretaceous to the Campanian,^[3][4] although Zhangheotheriidae might be paraphyletic in relation to other forms.^[4] Chronoperates has been suggested to be a late surviving representative of this clade,^[5] offering a ghost lineage extending to the late Paleocene; however, no recent phylogenetic studies have incorporated it.

Particular sub−groups of Symmetrodonta are better studied, e.g. Spalacotheriidae, which has acute−angled molariform teeth, strongly reduced talonids, and conspicuous anterior and posterior cingulids.

Though some forms like Zhangheotherium retain a Meckelian groove, at least Spalacotheriinae lost it, acquiring modern ear anatomy. Their deciduous canines and premolars as well as long lower jaw indicate a carnivorous/insectivorous diet.^[6]

Zhangheotherium was specialised to a tree-dwelling lifestyle.^[7] It shows evidence of tarsal spurs, indicating that, like most non-therian Mammaliaformes, at least some symmetrodonts were venomous like the modern platypus.^[8]

One species, Spalacotheridium noblei, is notable for its small size. It is one of the smallest known mammals. Each individual molar is little more than 0.25 mm across.

• Evolution of mammals

• Discovering a Missing Link

v t e Mammalia
Kingdom: Animalia Phylum: Chordata Clade: Synapsida Clade: Therapsida Clade: Cynodontia Clade: Mammaliaformes Synapsida see Synapsida Cynodontia see Cynodontia Mammalia see below↓
Mammalia Australosphenida? †Ambondro †Tendagurutherium? Monotremata? †Henosferidae Asfaltomylos Henosferus †Ausktribosphenidae Ausktribosphenos Bishops? Kryoparvus? Monotremata †Kollikodontidae? Kollikodon Kryoryctes? Sundrius †Steropodontidae Parvopalus? Steropodon †Teinolophidae Stirtodon? Teinolophos Ornithorhynchoidea †Opalios Ornithorhynchidae †Dharragarra? †Monotrematum? †Obdurodon Ornithorhynchus †Patagorhynchus? Tachyglossidae †Megalibgwilia †Murrayglossus Tachyglossus Zaglossus †Allotheria? see Allotheria includes multituberculates Theriimorpha see below↓
Theriimorpha Theriimorpha †Acinacodus †Amphidon †Amphilestes †Aploconodon †Chaoyangodens †Comodon †Condorodon †Dyskritodon? †Fruitafossor? †Hakusanodon †Indotriconodon †Juchilestes †Jueconodon †Kemchugia †Kryptotherium †Liaoconodon †Liaotherium †Manchurodon †Nakunodon †Phascolotherium †Sangarotherium †Tendagurodon †Allotheria? Monotremata? †Gobiconodontidae Fuxinoconodon Gobiconodon Hangjinia Huasteconodon? Meemannodon Repenomamus Spinolestes †Jeholodentidae Jeholodens Yanoconodon †Klameliidae? Ferganodon Klamelia †Triconodontidae Alticonodon Arundelconodon Astroconodon Corviconodon Eotriconodon Jugulator Meiconodon Priacodon Triconodon Trioracodon Victoriaconodon Volaticotheria? †Volaticotheria Argentoconodon Ichthyoconodon Triconolestes? Volaticotherium †Tinodontidae Gobiotheriodon? Tinodon Trishulotherium Yermakia Trechnotheria Monotremata? †Zhangheotheriidae Anebodon Kiyatherium Maotherium Ningchengodon Origolestes Zhangheotherium †Spalacotheriidae Akidolestes Aliaga Cifellitherium Heishanlestes Infernolestes Lactodens Shalbaatar Spalacolestes Spalacotheridium Spalacotherium Spalacotheroides Symmetrodontoides Symmetrolestes Yaverlestes Cladotheria see below↓
Cladotheria †Dryolestida Brancatherulum? Dorsetodon Drescheratherium Euthlastus Henkelotherium Paurodon Tathiodon Dryolestidae Achyrodon Amblotherium Anthracolestes Beckumia Crusafontia Dryolestes Groebertherium? Guimarotodus Hercynodon Krebsotherium Lakotalestes? Laolestes Minutolestes Phascolestes Portopinheirodon Treslagosodon †Meridiolestida Amarillodon Austrotriconodon Bondesius Casamiquelia Cronopio Groebertherium? Lakotalestes? Leonardus Necrolestes Paraungulatum Quirogatherium Solanutherium Mesungulatoidea Coloniatherium Mesungulatum Orretherium Peligrotherium Reigitherium †Donodontidae Amazighodon Anoualestes Donodon Stylodens Prototribosphenida †Abelodon †Amphibetulimus †Amphitherium †Arguimus †Brancatherulum? †Chunnelodon †Magnimus †Minimus †Mozomus †Nanolestes †Palaeoxonodon †Vincelestes Monotremata? †Peramuridae Afriquiamus? Kouriogenys Peramuroides Peramus Tendagurutherium? Tribosphenida / Boreosphenida †Aegialodon †Argaliatherium †Brasilestes †Carinalestes †Eomaia †Holoclemensia †Hypomylos †Juramaia †Kermackia †Kielantherium †Pappotherium †Picopsis †Tirotherium †Tribactonodon Theria †Patagomaia Eutheria see Eutheria includes placentals Metatheria see Metatheria includes marsupials
Other taxa Incertae sedis †Atlasodon †Chronoperates †Microderson †Thereuodon †Tingamarra Saint Bathans Mammal UA 8699 Other taxa Prototheria Yinotheria Eutriconodonta Amphilestidae Theriiformes Pantotheria Symmetrodonta Eupantotheria Paurodontidae Amphitheriidae Marsupionta