Dmitri Ivanovsky
Ivanovsky, c. 1915
Born	(1864-10-28)28 October 1864 Nizy, Gdovsky Uyezd, Saint Petersburg Governorate, Russian Empire
Died	20 June 1920(1920-06-20) (aged 55) Rostov-on-Don, Russian SFSR
Nationality	Russian
Alma mater	St. Petersburg Imperial University
Known for	Discovery of viruses, Tobacco mosaic virus
Scientific career
Fields	Virology
Institutions	St. Petersburg Imperial University Imperial University of Warsaw Saint Vladimir Imperial University of Kiev Donskoy University
Doctoral advisor	Andrei Famintsyn

Dmitri Iosifovich Ivanovsky (alternative spelling Dmitrii or Dmitry Iwanowski; Russian: Дми́трий Ио́сифович Ивано́вский;^[a] 28 October 1864 – 20 June 1920) was a Russian botanist, the co-discoverer of viruses (1892), and one of the founders of virology.^{[1][2][3][4][5]}

Ivanovsky was born in the village of Nizy, Gdov Uyezd. He studied at the University of Saint Petersburg under Andrei Famintsyn in 1887, when he was sent to Ukraine and Bessarabia to investigate a tobacco disease causing great damage to plantations located there at the time. Three years later, he was assigned to look into a similar disease occurrence of tobacco plants, this time raging in the Crimea region. He discovered that both incidents of disease were caused by an extremely minuscule infectious agent, capable of permeating porcelain Pasteur-Chamberland filters, something which bacteria could never do. He described his findings in an article (1892)^[6] and a dissertation (1902).^[7] Then he worked at the Imperial University of Warsaw and at Donskoy University in Rostov on Don.

In 1898, the Dutch microbiologist Martinus Beijerinck independently replicated Ivanovsky's experiments and became convinced that the filtered solution contained a new form of infectious agent, which he named virus. Beijerinck subsequently acknowledged Ivanovsky's priority of his discovery of the submicroscopic entity that was filterable.^[2]

• Lecoq, H (October 2001). "Découverte du premier virus, le virus de la mosaïque du tabac: 1892 ou 1898?" [Discovery of the first virus, the tobacco mosaic virus: 1892 or 1898?]. Comptes Rendus de l'Académie des Sciences. Series III – Sciences de la Vie (in French). 324 (10): 929–33. Bibcode:2001CRASG.324..929L. doi:10.1016/S0764-4469(01)01368-3. PMID 11570281.

• "Viruses and the Prokaryotic World". Retrieved 19 April 2008.

v t e Microbiology: Virus
History Introduction Social history of viruses Virology
Components	Capsid Viral envelope Viral protein
Viral life cycle	Viral entry Viral replication Viral shedding Viroplasm Virus latency Lytic cycle Lysogenic cycle
Genetics	Antigenic drift Antigenic shift Phenotype mixing Reassortment Viral evolution
By host	Animal virus Bacteriophage Virophage Human virome Mycovirus Plant virus
Other	Antiviral drug Giant virus Helper virus Viral vector Helper dependent virus Laboratory diagnosis of viral infections Marine viruses Neurotropic virus Oncovirus Satellites Viral disease Viral load Virus-like particle Virus classification Virus quantification Virome Virosphere
Portal Category Commons WikiProject

v t e Microorganisms
Groups	Archaea Bacteria Cyanobacteria Fungi Nanobacterium Prokaryote Protist Protozoa Virus
Microbiology	Microbial biogeography Microbial genetics Microbial intelligence Microbial metabolism Microbial phylogenetics Microbial population biology Mycology Virology
Motion	Bacterial motility run-and-tumble twitching gliding Protist locomotion amoeboids Bacteria collective motion
Ecology	Biofilm Biological pump Kill the Winner hypothesis Microbial consortium Microbial cooperation Microbial biodegradation Microbial ecology Microbial cyst Microbial food web microbial loop viral shunt Microbial mat Microbial synergy Microbiome microbiota holobiont Quorum sensing Host microbe interactions in Caenorhabditis elegans
Plants	Plant microbiome Root microbiome Seagrass microbiome Soil microbiology Spermosphere
Marine	Marine microorganisms Marine viruses Marine prokaryotes Marine protists Microalgae Antarctic microorganism Coral microbiome Hydrothermal vent microbial communities Marine microbial symbiosis Microbial oxidation of sulfur Phycosphere Picoeukaryote International Census of Marine Microbes
Human related	Microbes in human culture Microbiomes of the built environment Food microbiology Microbial oil Microbial symbiosis and immunity Nylon-eating Human microbiome asthma dysbiosis fecal gut lung mouth skin vagina in pregnancy placenta uterus Human Microbiome Project Protein production Synthetic microbial consortia
Techniques	Dark-field microscopy DNA sequencing Impedance microbiology Microbial cytology Microbial DNA barcoding Microbiological culture Staining
Other	Bioremediation Deep biosphere Microbial dark matter Microswimmer biohybrid Lines on the Antiquity of Microbes Microbially induced sedimentary structure Omics Physical factors affecting microbial life Siderophore
Category

Authority control databases
International	ISNI VIAF WorldCat
National	Germany Czech Republic Poland
Other	Encyclopedia of Modern Ukraine