A ctenidium is a respiratory organ or gill which is found in many molluscs. This structure exists in bivalves, cephalopods, Polyplacophorans (chitons), and in aquatic gastropods such as freshwater snails and marine snails.^[1] Some aquatic gastropods possess one ctenidium known as monopectinate and others have a pair of ctenidia known as bipectinate.

A ctenidium is shaped like a comb or a feather, with a central part from which many filaments or plate-like structures protrude, lined up in a row. It hangs into the mantle cavity and increases the area available for gas exchange.^[2] The word is Latinized but is derived from the Greek ktenidion which means "little comb", being a diminutive of the word kteis meaning comb.

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  Pair of large, grey gills visible inside the mantle cavity of a giant squid

v t e Cephalopod anatomy
Shell	Types External Ammonoid shell Argonautid eggcase Nautiloid shell Orthocone Internal Belemnoid guard Cirrate shell Cuttlebone Gladius (pen) Spirula shell Stylet Features External Aperture Apex Callus Lirae Periostracum Sculpture Spire Sutures Umbilicus Internal Body chamber Camerae Nacre Phragmocone Septa Siphuncle Whorls
Mantle & funnel	External anatomy Dermal structures Fins (wings) Flotation devices ("secondary fins") Funnel–mantle locking apparatus Tail Internal anatomy Branchial hearts Ctenidia (gills) Hepatopancreas (digestive gland) Ink sac and ink Needham's sac Nephridia ("kidneys") Nidamental glands Osphradium Pericardial glands
Head & limbs	Brachial crown Arms Hectocotylus Tentacles Carpus Dactylus Manus Suckers and hooks Buccal region Aptychus Beak Odontophore Radula Spadix and antispadix Occipital region Nuchal crest (occipital crest) Nuchal folds (occipital folds) Occipital membrane Olfactory organ Nuchal organ Other parts Eyes Statocysts
General	Chromatophores Photophores Nervous system Squid giant axon Squid giant synapse
Developmental stages: Spawn → Paralarva (Doratopsis stage) → Juvenile → Subadult → Adult • Egg fossils • Protoconch (embryonic shell)